Lawrence M. Witmer, PhD
Professor of Anatomy
Chang Professor of Paleontology

Dept. of Biomedical Sciences
Heritage College of Osteopathic Medicine
Life Science Building, Rm 123
Ohio University
Athens, Ohio 45701 USA




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Lawrence Witmer, PhD
1995 – present
webpage: University page
Ryan Ridgely, BS
Research Associate
2000 – present
Projects: NSF grant collaborator, µCT & 3D viz guru
webpage: University page
Wm. Ruger Porter, PhD
Lecturer, Postdoctoral researcher
Doctoral student: 2008 – 2015
Doctoral dissertation: Physiological Implications of Dinosaur Cephalic Vascular Systems.
webpage: University page
Current status: 2014 – present: Assistant Lecturer in Anatomical Sciences, OU Heritage College of Osteopathic Medicine
Don Cerio, PhD
Lecturer, Postdoctoral researcher
Doctoral student: 2012 – 2019
Doctoral dissertation project: The visual apparatus of avian dinosaurs and other diapsids: anatomical correlates of behavior and evolution..
Current status: 2018 – present Visiting Assistant Lecturer in Anatomical Sciences, OU Heritage College of Osteopathic Medicine
JP Nassif, BS
Doctoral student: 2015 – present
Doctoral dissertation project: Comparative morphology and biomechanics of the middle-ear conduction system in sauropsid reptiles.
webpage: Personal page
DJ Morgan, MS
Doctoral student: 2018 – present
Doctoral dissertation project: TBA
WitmerLab Alumni:  
Emily Caggiano, BS
WitmerLab research assistant: 2015 – 2019
current status: PhD student at MD Anderson UTHealth Graduate School of Biomedical Sciences, Houston, Texas
Catherine Early, PhD
Doctoral student: 2013 – 2019
Doctoral dissertation project: Quantitative assessments of avian endocasts as tools for inferring neuroanatomical traits and potential functional capabilities.
webpage: Personal page
current status: NSF postdoctoral fellow at the University of Florida

Ashley Morhardt, PhD
Doctoral student: 2010 – 2016
Doctoral dissertation project: Gross Anatomical Brain Region Approximation (GABRA): assessing brain size, structure, and evolution in extinct archosaurs.
webpage: Penchant4Paleo (blog),
current status: Assistant Professor of Anatomy, Washington University School of Medicine at St. Louis
Jason Bourke, PhD
PhD Recipient 2015
2008 – 2015
Doctoral dissertation project: Implications of airflow dynamics and soft-tissue reconstructions for the heat exchange potential of dinosaur nasal passages.
webpage: Personal page
current status; Assistant Professor of Anatomy, New York Institute of Technology's College of Osteopathic Medicine at Arkansas State University
Shayna Knece, BS
Master's candidate in the OU Patton College of Education
2013 – 2014
Masters thesis project: TBA.
Elsa Kendrick
Honors Tutorial College Thesis Researcher
2013 – 2014
Honors thesis project: Anatomy and evolution of the tongue and hyolingual apparatus in diapsids.
Current status: Bachelor of Science in Nursing 2019
Alexandra Spaw Johns, BS, MD
Honors Tutorial College Thesis Researcher
2012 – 2014
Honors thesis project: Fetal Developmental Anatomy of the Human Cardiovascular and Central Nervous Systems Using Lugol’s Iodine Staining and Micro-Computed Tomography
current status: Surgical resident at UC Davis
Eric Snively, PhD
2010 – 2013
Projects: Computational modeling of dinosaur head and neck function; Rigid body mechanics of prey capture in large carnivorous dinosaurs
webpage: UW-L page
current status: Assistant Professor, Biology Department, University of Wisconsin-La Crosse
Cheyenne Romick, BS
Honors undergraduate researcher
2011 – 2013
Honors thesis project: Ontogeny of the brain endocasts of ostriches (Aves: Struthio camelus), with implications for interpreting extinct dinosaur endocasts.
Joe Daniel, PhD
PhD Recipient 2012
2004 – 2012
Doctoral dissertation project: Heads and skulls as sediment sorters: an actualistic, CT-based study in taphonomy.
webpage: Paleoaerie
current status: Gross Anatomy Instructor at the University of Arkansas for Medical Sciences
Dave Dufeau, PhD
PhD Recipient 2011
2005 – 2011
Doctoral dissertation project: The evolution of cranial pneumaticity in Archosauria: patterns of paratympanic sinus development
webpage: University page
current status:  Assistant Professor of Anatomy at Marian University College of Osteopathic Medicine, Indianapolis, IN
Amy Martiny, MA
MA recipient 2010
2008 – 2010
Project: sociology of science and medicine, photography and web delivery of WitmerLab cast collection
current status: Surgical Skills Lab Specialist at the Department of Neurological Surgery at Northwestern's Feinberg School of Medicine, Chicago.
Tobin Hieronymus, PhD
PhD Recipient 2009
2002 – 2009
Doctoral dissertation project: Osteological correlates of cephalic skin structures in Amniota: documenting the evolution of display and feeding structures with fossil data.
webpage: University page
current status:  Assistant Professor, NEOMED
Kate Burns
High School Intern from Linworth Alternative Program, Columbus, Ohio
2 Feb – 26 Mar 2009
Projects: 3D viz of crow brain & inner ear, painting casts of T. rex, Gorgosaurus, and Diplodocus, specimen curation
current status: Ohio University graduate
Takanobu Tsuihiji, PhD
Postdoctoral Fellow
2006 – 2008
Projects: Cervicocephalic anatomy of archosaurs, Mongolian dinosaurs
current status: Associate Professor at University of Tokyo, Japan
Justin Tickhill, MS
Masters recipient 2007
2004 – 2008
Masters thesis project: The Virtual Pig Head: digital imaging of cephalic anatomy.
current status: Gross Anatomy Instructor, Grand Canyon University, Phoenix AZ,
Casey Holliday, PhD
PhD recipient 2006
2000 – 2006
Doctoral dissertation project: Evolution and function of the jaw musculature and adductor chamber of archosaurs (crocodilians, dinosaurs, and birds).
webpage: Holliday Lab
current status: Associate Professor, Department of Pathology and Anatomical Sciences, University of Missouri
Brian Beatty, PhD
PhD student
webpage: The Aquatic Amniote Blog, University page
current status: Assistant Professor, Department of Anatomy, New York College of Osteopathic Medicine
Andy Clifford, PhD
BS Honors 2000; MS recipient 2003
1999 – 2000, 2001 – 2003
Honors thesis project: The narial anatomy of hooded seal (Cystophora cristata) with respect to other Carnivora.
Masters thesis project: Narial novelty in mammals: case studies and rules of construction.
current status: Formerly Brown University, PhD student; current status: Hathaway Brown School, Cleveland, OH.
Pat O'Connor, PhD
PhD recipient 2003 (State University of New York at Stony Brook under C. A. Forster, Witmer committee member)
2000 – 2002 (completed doctoral research in the WitmerLab while serving as an anatomy instructor)
Dissertation title: Pulmonary pneumaticity in extant birds and extinct archosaurs.
webpage: University page
current status: Professor, Department of Biomedical Sciences, Ohio University Heritage College of Osteopathic Medicine
Jayc Sedlmayr, PhD
PhD recipient 2002
1997 – 2002
Dissertation title: Anatomy, evolution, and functional significance of cephalic vasculature in Archosauria.
webpage: University page
current status: Assistant Professor, Department of Cell Biology and Anatomy, LSU Health Sciences Center
Mike Papp, MS
MS recipient 2000
1996 – 2000
Thesis title: A Critical Appraisal of Buccal Soft-Tissue Anatomy in Ornithischian Dinosaurs
current status: Project Geologist, AECOM, Hebron, KY
Thomas Carr, PhD
PhD student
1996 – 1997
Projects: tyrannosaur skull structure, analysis of CT data of sauropod dinosaurs
webpage:University page
current status: Associate Professor of Biology, Carthage College, Kenosha, WI
Click the image at right to see the archived WitmerLab People page.

Last updated: 08/21/2019